Prologue of the film depicts Jim Hawkins as three years (voiced by Austin Majors) reading a storybook to read. Jim is enchanted by the tales of the legendary pirate Captain Flint and his ability to appear from nowhere, raid passing ships, and disappear, to hide the loot on the mysterious "Treasure Planet". Twelve years later, Jim (now the voice of Joseph Gordon-Levitt) has become a distant and alienated teenager. He is shown begrudgingly helping his mother Sarah (Laurie Metcalf) run an inn and deriving amusement from "solar surfing" (a hybrid of skysurfing and windsurfing atop a board attached to a rocket powered by solar energy), a pastime he often gets him into trouble.
One day, a spaceship crashes near the inn. The pilot dies, Billy Bones (Patrick McGoohan), gives Jim a sphere and tells him to "beware the cyborg". Shortly after, a band of pirates plunder and burn the inn. Jim, his mother, and their dog-like friend Dr. Delbert Doppler (David Hyde Pierce) barely escape. The sphere turns out to be a holographic projector, showing a map that Jim realizes leads to Treasure Planet.Doppler commissions a ship called the RLS Legacy, on a mission to find Treasure Planet. The ship is commanded by the cat-like, quick-witted Captain Amelia (Emma Thompson), together with his partner from the skin of stone and covered in the first place, Mr. Arrow (Roscoe Lee Browne). The crew is a diverse group, secretly led by cook John Silver (Brian Murray), who Jim suspects is the cyborg he was warned. Jim is sent to work in the kitchen, despite his mistrust of Silver, soon forming a tenuous relationship between father and son (a montage with the song "I'm Still Here" shows Jim and the bond cyborg on various sailing chores, interspersed with childhood flashbacks of Jim, during which his father appears indifferent to him, and finally leaves without warning when Jim is a pre-teen). During a meeting with a supernova, Silver falls in water but is rescued by Jim. The supernova then devolved into a black hole, where Arrow is adrift at sea and is lost, so Jim is criticized for failing to ensure the lines of life, while, in fact, Arrow has been cut by a crew member insectoid ruthless named Scroop (Michael Wincott).
As the ship reaches Treasure Planet, mutiny erupts, led by Silver. Jim, Doppler and Amelia abandon ship, leaving the map behind unintentionally. Silver, who believes that Jim has the map, has the opportunity to kill Jim, but refuses to do so because of his attachment to the boy. The escapees were shot down by a mutineer during their escape, causing injury to Amelia.
While exploring Treasure Planet Forests, the fugitives meet BEN (Martin Short), an abandoned robot extravagant claims to have lost much of his memory and invites them to his house to treat the wounded Amelia. The pirates corner the group here: using a back-door, Jim and BEN return to the ship in an attempt to retrieve the map. Scroop, aboard the ship as lookout, stalks and fights Jim. BEN, working to sabotage the ship's artillery, accidentally turns off the artificial gravity, after Jim and Scroop threaten to float in space. Jim takes the pole, while Scroop becomes entangled in the flag and cut free, while Scroop floats away, presumably to his death. Jim and B.E.N. obtain the map. On their return, they are captured by Silver, who has already captured Doppler and Amelia.
When Jim is forced to use the map, the group finds their way to a portal that can be opened anywhere in the universe, this being the means by which Flint conducted his raids. The treasure is at the center of the planet, accessible only via the portal. Treasure Planet is proving to be a large space station built by unknown architects and seconded by Flint. In light of the Treasury, Jim meets the skeletal remains of Flint itself, with a missing piece of computer cognitive Ben. Jim replaces this piece, causing B.E.N. to remember that the planet is about to explode at the time of the discovery of the treasure. In the ensuing catastrophe, Silver finds himself torn between holding on a literal boat load of gold and saving Jim, hanging from a cliff after a fall. Silver saves Jim, and the group escapes to the Legacy, which is corrupt and lacks the driving force necessary to leave the planet in time to escape. Jim attaches a rocket to a thin plate of metal and rides to the portal to open in a new location, while Doppler pilots the ship behind him. Jim manages to open the portal to his world of origin spaceport, through which all fleeing the destruction of Treasure Planet.
After the flight, Amelia has the surviving pirates imprisoned aboard the ship and offered to recommend Jim Interstellar Academy for his heroic actions. Silver sneaks below deck, where Jim finds him preparing his escape. Jim lets him go, inheriting Silver shape-changing pet called Morph (Dane A. Davis). Silver expected to Jim "to shake the stars", then throws a handful of jewels and gold he had taken from Treasure Planet to pay for the reconstruction of the inn. The film ends with a party at the rebuilt inn, showing Doppler and Amelia now married with children, and Jim a military cadet. He looks to the sky and sees the image of Silver in the clouds.
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